She was believed to be around 141 years old when she was euthanized due to "ongoing bone conditions related to advanced age," the zoo said Known as "the Queen of the Zoo," Gramma was in the first ...

Gramma, the beloved Galapagos tortoise who lived at the San Diego Zoo for nearly a century, has died on Nov. 20. She was (about) 141. The zoo's care team "made the compassionate and exceptionally ...

Gramma, a 141-year-old Galapagos tortoise and the San Diego Zoo’s oldest resident, has died. She came from the Bronx Zoo nearly a century ago and was known as the "Queen of the Zoo." Visitors and ...

Gramma, a Galápagos tortoise that the San Diego Zoo called “a quiet and constant presence” for nearly a century at the Zoo, has died. Yes, a century, dating back to the zoo’s early days. The zoo, in a ...

Gramma, the beloved Galapagos tortoise who lived at the San Diego Zoo for nearly a century, died Thursday. She was about 141. The zoo’s care team “made the compassionate and exceptionally difficult ...

After more than a century of munching on her favorite foods of romaine lettuce and cactus fruit, beloved Galápagos tortoise Gramma, the oldest resident of the San Diego Zoo, has died. Gramma was born ...

LOS ANGELES -- After more than a century of munching on her favorite foods of romaine lettuce and cactus fruit, beloved Galapagos tortoise Gramma, the oldest resident of the San Diego Zoo, has died.

Gramma the Galapagos tortoise was a roughly 141-year-old icon for visitors to the San Diego Zoo. After nearly a century of delighting guests, the zoo shared on social media this week that she died ...